Seiichi Morimura, who wrote a searing expose of the Japanese army’s secret biological warfare program in occupied China, describing how it forcibly infected thousands of prisoners with deadly pathogens, died July 24 in Tokyo. He was 90.

The announcement of his death by his publisher, Kadokawa, was cited in Japanese media.

Morimura detailed the atrocities committed by the Japanese program — called Unit 731 — in a widely sold book, "Akuma no Hoshoku,” or "The Devil’s Gluttony” (1981). Among the horrors he described were vivisections performed without anesthesia on those who had been deliberately administered germs; doctors wanted to see firsthand how the ensuing diseases infected the body.